OMG this game was amazing. I loved finding secrets everywhere, and the artstyle was really nice. The movement felt good aswell (I loved dashing around like a maniac). The sound was also really good. However, I did found some small things that you could improve on.
For example the spike traps. The spikes made a sound when they should release. But I feel like this sound came to early. The sound came before the spikes moved, maybe you wanted this to be an audio clue so you know when the spikes are going to come. But if you want to have it so that the sound is when the spikes come out I think you need to redo it. Maybe it works fine for you but not for me because we have different fps?
Another thing was when you killed a boss. The meter was still displayed until you went out of that section of the map. You can see it on the image below.
Another thing I found when I met one boss was a place to stay to kill the boss without getting damaged. You can see the position and which boss on image below.
It seems like the brush upgrades stacks. At first I thought this was really cool. I felt amazing walking around with the biggest brush ever. Then I thought it might be too op(I killed a whole room of enemies without seeing one). But then I remember the spider enemies(I hated those spiders). If they shoot at you when you are that big it is incredible hard to dodge the webs because the brush takes up so much space. I did never encounter a spider when I was this big but I believe that it would be hard to defeat them. On the image below you can see what I mean with the big brush. I think I had three upgrades here.
I found another bug/glitch in the beginning of the game. I had just died from a boss and was angry so I dashed upwards in frustration. I lost 4 hearts of this. But when I got to the next room I had full health. Maybe the game can't keep up with my fast dashes :D Here is a link to a video that shows what happened. https://streamable.com/14osw .
Well that was all minor bugs/glitches I found. I would totally buy this game if you release a bigger version of it! You should put it on steam for 10-20 dollars. Thank you for the gamejam btw!
Edit: I played this game in broswer. Was there a reason to have a setup instead of just a zip? My antivirus program did not let my play the downloaded version.