Wow, i now have replayed the final-version, and i put my ratting up accordingly. For me, this is by far the hottest & cuttest game of this jam. The little brushing sound you had is just perfect and had to the "tactile" feeling, the color-effects works really well, and the "apex and after" feel strong & accurate.
For me, the straightforwardness of your game is a great force : everybody colored something some day, we all know how to do it. The only way i think you could improve it would be adding more and more little things, like specific reaction when you colorize specific parts of the critter, a more polished interface, and so on.
Whatever you do with it in the futur : great work.