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Great take on the theme. Mushrooms were fun to find since they spawned in random locations each game across the map.  Unlike some of the others I didn't feel annoyed since only the keys on the same axis were swapping in my first game. However when I played it a second time, I understood what they were talking about. The confusing part was that I would go forward with D and rotate with S. But I think the confusing controls would really balance out in local multiplayer, the intended way of playing leading to a hectic yet enjoyable experience.

Yes. That was the ultimate goal. When two people are annoyed by a competitive game together, it comes out to be a real nice local multiplayer session and that was the intention for me with this.  Actually the concept that I had in mind was that red animals will rotate your key controls clockwise and blue ones would do anti-clockwise so once you figure this out, instead of getting hit by the red animal thrice again to get back to normal, you can find the path of a blue animal and get hit by that once and save some time in the local multiplayer game. Just an added optional strategy. But I think there is a bug in the collision where it rotates twice and hence you get inverse controls on the same axis. That technically should have happened after two collisions.