Wow, this is great! Super polished and generally well executed. I like that you experimented with a variety of mechanics and then played them off one another.
I think there's a tension between the more puzzle-y side of the game and the dexterity platforming side which you did a good job balancing. The one mechanic that maybe leaned too far in the dexterity direction is the controls reversal. That mechanic felt more annoying than interesting to me. But maybe that's just because I have no dexterity, but like puzzles :)
Ignoring that one nitpick, your game was really fun! Great job overall!