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Does this same creature design consistently glitch out in the current released version (other than Android)? If so, then the glitch wasn't introduced in the Optimization branch. There were definitely glitches happening before but they definitely increased drastically after my optimization attempts. 

I'm in the process of adding import and export functionality to both creature designs and simulation save files. I'll let you know once that's ready so you can send me this design to test with.

Yes, it does glitch out in both 2.0.3 and 2.0.4 (Windows x64)

Thanks, I'll send you a small file of the simulation run, I believe you can get the creature from there? Two files in fact, the first is of the creature before some slight changes which made the glitches significantly worse. Maybe these changes (bone length if I recall) will give a hint.