I enjoyed playing this quite a bit. A try and retry that rewards you for playing it for a while.
There were a few times per run when the scene would "skip/skid" upwards, which is most likely a bug. I also had a bit of a hard time focusing in on what was an obstacle vs what was just part of the scene. I feel that this could have been avoided with better color palette choices. The placement of the UI (especially Options) was a little underthought, but loved what I saw once I opened it up.
At first, I couldn't quite tell why I wasn't able to level up the different upgrades, but eventually realized that there was something that I was picking up. I still don't know what that exactly was (where am I getting the scales and fire? from the stuff that I'm supposed to be avoiding? How does that work?). I most likely missed something, so do let me know what it was!
Great job putting this together, definitely a nice entry this time around!