Your question was clear... But so was the answer, I'm afraid.
If you would like it more technically worded:
- The 3d assets he used to design the characters are included in the game Honey Select (used with permission) - or possibly its marketplace.
- The tool he used to combine those assets, pose the characters, and export them into static images and animations is the game Honey Select.
- The ultimate source of the characters' *design* is Runey's own imagination - as limited by the assets available to him due to the toolset he is using.
I hope that answers the question, otherwise I am not sure what you are asking.
Worst case scenario, try asking in (I assume) French.