My thoughts on your game:
- Animations: the walk animations are inverted and you should check if the feet are touching the ground to play it. Maybe they were too detailed, but looked cool anyways. I am not and expert on sprite animation, but you can take a look at the video to check what I mean with "too detailed".
- Mechanic: the upgrades were a bit difficult to interpret, you could have written some pieces of this awesome hand-written text that you used for the tutorial (I liked it, it was quite funny).
- The collision shape of the player is too big. Unless you are doing a rage pixel-perfect game, the collider of the player should usually be smaller that the sprite, to make it a bit snappier.
I liked your all over the place story-telling, and the wall running was good.