Very interesting main mechanic - it is definitely quite unique and fun to explore, could be made into a full fledged game if designed properly. The overall atmosphere and visuals help it too.
Im not really sure if im a fan of the following trail mechanic. Feels like it doesnt serve much of a purpose, if we want to punish the player for not moving and visiting same spot too much, it would need to be either just health damage, or actually full on focus on this mechanic, and ideally give the player a small time rewind or something, as it only feels frustrating, makes you replay the whole level and feels like it did not even have to be there IMO (i know it represents one of the jam themes, so i would go with the damage option)
The game feels a bit clunky or glitchy at times - the boar does not move as soon i get in, and feels stuck. The abilities do not feel responsive, eg. the climbing sometimes work sometimes not really, etc
Other than that the game is actually nice, it is definitely quite a big one for a game jam, but i think you managed quite well. Was very enjoyable to play, thank you :)