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Hi, 

I love your program, I am really happy that you have made such an amazing tool. 

I was wondering if there is a way to have the pngtuber to take desktop audio as microphone input to react to. 

Glad you're happy with it! 

Depending on your sound card, there should be a microphone device that you can enable on your PC's sound settings.  Mine is called "Stereo Mix".