This game was a lot of fun! Really fun and interesting gameplay mechanics. The boss battles had a good bit in difficulty and I love it. Almost everything in this game is really really good.
The only bad detail that I had with this are how blurry the character icons are during cutscenes.
I have a suggestion for another boss idea (if you ever think about making more bosses): A boss that can switch directions and also has the same abilities as Skeeter.