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The problem is that dot files (._000atl.rpy) which are metadata files that appear (normally if you access the files from a non mac network share iirc) were in the directory.  To fix this I had to:

dot_clean [App Bundle File]
xattr -cr [App Bundle File]

If you do these steps before you zip the application it should avoid this problem. Simply moving the file (and I know why you suggested it) isn't enough

As far as I know, ._ files are temp files generated by macOS, and aren't actually present in the game files when I build the mac version, nor after it's built, so there isn't much I can do about it unfortunately. Still, I'm glad you were able to find a solution.