The shotgun is way too powerful compared to the other weapons. I think it's because the bugs are set to disappear based on the number of hits they take. It would've been even better if the single-shot rounds had piercing capabilities.
The ground bugs felt a bit trivial because of the tank's main cannon and jump attacks. However, the aerial bugs were such relentless bullet sponges that I eventually got overwhelmed and defeated. Also, I didn't really see a reason to use the right-click dash toward the late game.
That said, my second playthrough was still very satisfying. The controls feel amazing!