Really good game! I really liked how you contextualize the abilities, and the world is very charming. I liked all the different areas you added.
Some small stuff, dying on one hit on the spikes felt a little bit too punishing, and on the heart boss you would jump up to hit it and at the same time a spike spawn on top of you so it would be really hard to react (maybe having some effect that shows when a spike is about to fall could help?)