Thanks for playing and for the detailed feedback.
The organic zone is definitely the worst designed zone imo. It's the first zone I made and I don't have a ton of experience making metroidvanias so I was learning a lot about area design during this jam (the only other metroidvania I've made was for MVM 26 and it was a complete dumpster fire). I wanted to do a full redesign, but only had time for some minor adjustments at the end of the jam. But since most of the tracks in the game are loops you should be able to focus on defeating all the enemies in a room first, then find somewhere safe to look around (eg, the first room in the organic zone has a track with a save point and no enemy bubbles, which is also a pretty decent look-out spot). Aiming with a controller also sucks, which is why there were so many warnings. In hindsight I probably should have made some kind of aim assist system for people using controllers.
If you don't mind me asking, what elements specifically confused you? There aren't a lot of objects in the game, and the only one that's meant to be somewhat mysterious are the beacons. But I'm also not very good at world design since most of the game's I've made are smaller arcade style games that don't have much to figure out. Thanks.