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I have a question for everyone who has already asked themselves this question. In the last H-scene with Ryoko you can see how the queen wants to renew Ryoko's chain but that would normally mean that the queen must be Ryoko's mother since she gave her the chain back then and only the person can remove the chain etc. but we have seen both characters and they don't look alike there aren't even any similar features why do you think the queen was able to renew Ryoko's chain anyway.

(+1)

Option 1, wild recessive genetics

Option 2, there is a secret why the chain restrictions dont fully apply here

Im leaning towards 2