I. Love. Incremental. Upgrades! I especially love it when incremental upgrades are uncapped! By the end of this game, I was zipping around with unlimited dashes and very high movement speed, and I always love movement speed. It's awesome! There were a couple issues I had so here they are.
1. Boss upgrades don't carry over when you die. Normally you keep all of your upgrades, so I assumed the same would apply to the major boss upgrades. Unfortunately, they don't, and there's no indicator I noticed that you lost them. I was confused why I couldn't get out of the second level until I returned to the boss room and saw they were back.
2. When you die, the amount of xp thingies you need to evolve resets to the starting amount, which might be intentional, but it just makes it so dying quickly becomes the optimal course of action so you can grind out evolutions.
3. The feedback for damaging an enemy seemed a bit inconsistent. Maybe it's because I'm an insane speed demon so my damage upgrades were a bit lacking, but it felt like the spores didn't work at all, and my lack of control over my non-dashing attacks just felt a bit sluggish, but again, might just be an upgrade path thing.
And once again someone's first game jam game is amazing. This is a great game, good job, I hope you continue to make more!