Hello! :)
In the jam version, it took me some time to get the movement/attack system. I believe it took some minutes to understand that the attack pattern was based on the movement direction of the unit. But after some time I got the hang of it and it was fun. :)
However, since you are making a more complex version now, it seems that a simpler move/attack system is a good choice. Also, you comparison to chest pieces makes it almost instantly understandable.
Being able to change runes every day and, later, even changing between two on the same day also seems cool, giving more possibilities, maybe even using one 'set' during the day and another at night.
In short, great changes! Congrats! :D