Nice game man, I still remember the game jam version!
It was fun to play and I love the additions you made to the game. The speed + damage upgrade path is still the way to go for me, and made it quite easy to just click and hold.
I then tried to just focus on health-regen and was able to tank so much damage haha.
I'd add a black overlay to the background when the Upgrade panel is opened, since I kept trying to click the next wave button when the panel was opened.
Some waves (like the small zombie waves which occurred quite often) gave barely any gold which was quite annoying at times.
Fun game though, but it became a bit to easy after a couple of waves