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Great work, it's even more impressive knowing you took on so many roles! It reminds me a little bit of a game I'd see on Nitrome (which is very high praise). I agree with WaffleExploder. Those rusty sign posts you have in a few levels could be a good place to add helpful info on controls and backstory. 

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Adding tutorials to those signs was the reason I made them, but since I was so ambitious with the game I sadly didn't have time to implement it. Thanks for playing!