Fun game -- I played to the end and enjoyed it. Simple but effective art, nice contrast in the palette. I also re-keyed to a two-handed control system as others did, and the keybinding works great. Great job on this!
I would suggest defaulting to having the sound effects on, as I nearly gave up at the first puzzle because there were no sound cues and I didn't realize I had to turn them on. There were also a few spots where the shots were firing so quickly that I think passing through was a matter of luck -- I would suggest reducing those, as it's not a great feeling when an outcome isn't in your control, even for people who love frustratingly difficult platformers (like me). Overall great work!