I've compiled the model and got the same error, I don't know if it's a bug with the program because your model seems perfectly fine, my best guess was that there's something wrong with the rolls but the program is designed so that's not a problem.
I tried using the repose button which the model already is and fixed a few rotations and weirdly enough that seemed to fix the problem for the most part which I'm not sure why, I'll try to see if this is a bug or not