Just came back to this today, love the new additions! The new balancing made the first few layers feel much less grindy, and the new upgrades really helped! I also liked the new songs :)
The menu redesign was super nice, and felt much more intuitive also, and I really appreciate you taking our feedback and adding (un)inverted controls!
Though there was one issue that made things almost unplayable until I figured out what was wrong: the movement is WAY too fast - it was to the point where if I tapped a movement key the entire tree would be off my screen in just 1 frame, and I thought it was broken. In the past I had used custom speed 20, but now I had to set it all the way down to 0.1 to get a similar speed. Should probably fix this ASAP.
Btw, I managed to find the song I was looking for, it was the "tires on fire" by coda, thanks for helping me out with that! also the credits was a nice touch for anyone else who may want to find a song. You might want to update it with credits for the new songs too.
But yeah, overall, great stuff!