Another thought I had. Their whole civilization, the entire species, they had to have had previous human contact somehow? Dog breeds didn't and wouldn't occur naturally via evolution. They required humans getting there grubby fingers into their DNA via selective breeding to get even just the breeds in the story. If not, they would've ended up more or less like Adastra. A wolven population with only things like color, size, build etc, not dobbies, Saint Bernards, and whatever Rhoun is.