Copy/pasting from the game's main page here:
Controls:
WASD - movement
Shift - sprint
Space bar - dodge
Left click - left weapon attack
Right click - right weapon attack
Scroll up - switch left weapon
Scroll down - switch right weapon
R - heal
F - interact
Esc - pause menu (WARNING: Enemies won't wait for you to unpause)
Ammo held is limited by the gun's magazine size as there is no reloading, so use your ammo wisely! Picking up certain guns will refill their respective ammo types.
Progress through the map by picking up keys to unlock new areas behind energy barriers until the egg is secured!