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Art:

- Gamma is a bit dark, maybe bumping up the light would help navigation.

- Essences left behind by the creatures to suggest prior habitation.

Design:

- Positioning trees to guide the player subliminally down an intended path.

- Maybe add elements throughout the game that are somewhat mysterious and unexplained to elicit the feeling of unknown and supernatural.

Gameplay:

- Creatures seem to teleport/move quickly to the player once past a certain location. I suggest lowering their speed and allowing them to be introduced from afar, then maybe adjusting their mechanics to spawn and attract randomly thereafter.